Digestive Systems of Animals

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The simplest invertebrates (animals without backbones) do not have specialized digestive organs. Single-celled organisms, such as amoebas, rely on intracellular digestion (digestion within the cell). Some many-celled organisms, such as the sponge, also use intracellular digestion. The sponge obtains the tiny organic particles that make up its diet from water passing through its body. Water enters through the sponge’s pores and leaves through an opening called the osculum. As water flows through the interior canals of the sponge, specialized cells that line these canals, called collar cells, catch and engulf organic matter. Inside the collar cells, sacs called vacuoles form around the food and enzymes digest it. The digested food then passes to other cells in the sponge’s body.

Intracellular digestion meets the needs of simple animals, but more complex organisms require systems that are more specialized. Animals such as jellyfish and nonparasitic flatworms combine the intracellular process with some specialized digestive organs. These animals have a definite mouth and a saclike cavity, which is lined with digestive cells that secrete enzymes. Digestion begins when the enzymes break down food inside the cavity in an extracellular (outside the cell) process. Cells then engulf the partly digested food, and an intracellular process similar to that of sponges completes digestion. Wastes are excreted through the mouth.

Most of the more complex invertebrates and all vertebrates (animals with a backbone) digest food entirely through extracellular processes. Food moves in one direction, from mouth to anus, through the series of organs that make up the alimentary canal. Specialization of various parts of the alimentary canal improves the body’s ability to break down food and absorb various kinds of nutrients. The mouth of many animals contains teeth or other structures to break up large lumps of food. Behind the mouth, the pharynx and esophagus swallow the food and move it to the stomach. The stomach temporarily stores the food, mixes it with digestive juices, and carries out some digestion.

Digestion is completed in the intestine. The liver and pancreas pour their digestive juices into the anterior (front) end of this organ. After the anterior intestine absorbs the usable products of digestion, the walls of the posterior (rear) intestine absorb leftover water. In vertebrates the anterior intestine is called the small intestine; the posterior intestine is the large intestine. Feces, composed of unabsorbed and indigestible food residues, form in the posterior intestine, where they are stored until they are excreted through the anus.

Within this basic plan, the specific components of the digestive system vary enormously from one animal to another. For example, a fish’s pharynx contains gill slits for breathing but has no digestive function. An earthworm’s stomach consists of two organs: a crop, in which food is stored, and a muscular gizzard, which carries out mechanical digestion by grinding food against particles of sand. The stomachs of ruminant mammals, such as cattle and deer, consist of three or four compartments, each performing a specific function. Amphibians, reptiles, and birds have an organ called a cloaca, which serves as an exit for both digestive wastes and sex cells.

Circulatory Systems in Non-Humans

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One-celled organisms and many simple multicelled animals, such as sponges, jellyfishes, sea anemones, flatworms, and roundworms, do not have a circulatory system. All of their cells are able to absorb nutrients, exchange gases, and expel wastes through direct contact with either the outside or with a central cavity that serves as a digestive tract.

More complex invertebrates have a wide range of circulatory system designs. These invertebrate circulatory systems are classified as either open or closed. Open systems—found in starfishes, clams, oysters, snails, crabs, insects, spiders, and centipedes—lack capillaries, and the blood bathes the tissues directly. In closed systems, the blood is confined to a system of blood vessels. Invertebrates with closed systems include segmented worms, squids, and octopuses.

All vertebrate animals have closed circulatory systems. These systems are classified by the number of chambers in the heart, which determines the basic configuration of blood flow. Fish have two-chambered hearts with one atrium and one ventricle. Blood pumped from the ventricle travels through arteries to the gills, where it diverges into capillaries and exchanges gases. Leaving the gills, the capillaries reconvene into blood vessels that carry the oxygenated blood to the rest of the body, where the vessels again diverge into capillaries before reconvening into veins that return to the heart. In this way, the blood passes through first the respiratory organs (the gills) and then the systemic circulation between each pass through the heart.

Frogs and amphibians have three-chambered hearts, with two atriums and one ventricle. Blood pumped from the ventricle enters a forked artery. One fork, the pulmonary circulation, leads to the lung. The other fork, the systemic circulation, leads to the rest of the body. Blood returning from the pulmonary circulation enters the left atrium, while blood from the systemic circulation enters the right atrium. Although there is some mixing of oxygenated and deoxygenated blood in the ventricle, a ridge within the ventricle assures that most of the oxygenated blood is diverted to the systemic circulation and most of the deoxygenated blood goes to the pulmonary circulation. In reptiles, this ridge is more developed, forming a partial wall. In crocodiles, the wall is complete, forming a four-chambered heart like that found in mammals and birds.

Aquaculture

Aquaculture, farming of aquatic organisms in fresh, brackish or salt water. A wide variety of aquatic organisms are produced through aquaculture, including fishes, crustaceans, mollusks, algae, and aquatic plants. Unlike capture fisheries, aquaculture requires deliberate human intervention in the organisms' productivity and results in yields that exceed those from the natural environment alone. Stocking water with seed (juvenile organisms), fertilizing the water, feeding the organisms, and maintaining water quality are common examples of such intervention.

Most aquacultural crops are destined for human consumption. However, aquaculture also produces bait fishes, ornamental or aquarium fishes, aquatic animals used to augment natural populations for capture and sport fisheries (see Fishing), algae used for chemical extraction, and pearl oysters and mussels, among others.

Aquaculture is considered an agricultural activity, despite the many differences between aquaculture and terrestrial agriculture. Aquaculture mainly produces protein crops, while starchy staple crops are the primary products of terrestrial agriculture. In addition, terrestrial animal waste can be disposed of off-site, whereas in aquaculture such waste accumulates in the culture environment. Consequently, aquaculturists must carefully manage their production units to ensure that water quality does not deteriorate and become stressful to the culture organisms.

Ruminant

Ruminant, even-toed animal that regurgitates and masticates its food after swallowing. The majority of ruminants have four nipples; they usually have sweat glands only on the muzzle and between the toes. Most species bear horns that may be permanent or may be shed periodically. The division includes three subdivisions: Tragulina, containing the chevrotains and characterized by a stomach with three chambers; Tylopoda, consisting of the camel, dromedary, llama, alpaca, guanaco, and vicuña, and characterized by a stomach with three distinct chambers; and Pecora, containing all sheep, goats, antelope, deer, gazelles, giraffes, and domestic cattle, and characterized by the presence of a distinct four-chambered stomach. Pecoran animals are known as true ruminants. Between the esophagus and the intestine, the stomach chambers of a true ruminant are the rumen, the reticulum, the omasum, and the abomasum, or rennet bag.

Scientific classification: Ruminants make up the suborder Ruminantia of the order Artiodactyla.

Amoeba

Amoeba, any of a group of unicellular organisms characterized by their locomotive method of extending cytoplasm outward to form pseudopodia (false feet). The amoeboid group includes hundreds of different organisms, ranging in size from about .25 to 2.5 mm (about 0.0098 to 0.098 in). Amoebas are considered the most primitive animals and are classified in the kingdom Protista. All amoeboid organisms have thin cell membranes, a semirigid layer of ectoplasm, a granular, jellylike endoplasm, and an oval nucleus. Some species live on aquatic plants and some in moist ground; others are parasitic in animals.

Amoebas also use pseudopodia for feeding. Chemical stimuli from smaller organisms, the amoeba's food, induce the formation of pseudopodia, pairs of which envelop the organism, at the same time forming a cavity, or vacuole. A digestive enzyme secreted into the cavity breaks down this food into soluble chemical substances that then diffuse from the cavity into the cytoplasm. Undigested food and wastes are excreted through the ectoplasm, which also absorbs oxygen from the surrounding water and eliminates carbon dioxide, a by-product of metabolism, in a form of respiration. After a period of growth, the amoeba reproduces by splitting into two equal parts.

At least six forms of amoeba are parasitic in humans. Most important of these is Entamoeba histolytica, which causes amebiasis and dysentery. The diseases often occur in epidemics when raw sewage contaminates water supplies or when soil is fertilized with untreated human wastes.